Captain Michael Clarke said: "It was a tough decision. They're two very good wicketkeepers and whichever way we went, the team was going to be fine."
The first of the three-Test series is in Brisbane, starting on 9 November.
The other notable omission from the squad was 19-year-old fast bowler Pat Cummins, who claimed seven wickets in his only Test appearance - against South Africa in Johannesburg last year.
"We didn't feel as though Pat Cummins was ready to play in a Test match," said chief selector John Inverarity.
The 12-man squad includes four pace bowlers - Ben Hilfenhaus, Peter Siddle, James Pattinson and Mitchell Starc - and spinner Nathan Lyon.
Australia squad: Michael Clarke (captain), David Warner, Ed Cowan, Shane Watson, Ricky Ponting, Mike Hussey, Matthew Wade (wk), Peter Siddle, Ben Hilfenhaus, Mitchell Starc, James Pattinson, Nathan Lyon.
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